Developmental patterns of Dracocephalum fragile Turcz. ex Benth. (Lamiaceae) in Tuva

We studied the developmental patterns of longrhizomatous-taproot herbaceous polycarpic of Dracocephalum fragile Turcz. ex Bentham individuals from Tuva. The base of the shoot system of adults is made up of polycyclic longrhizomatous-middlerosulate monocarpic shoot. A life form and features of shoot formation in ontogeny are described. Similarities and differences of ontomorphogenesis of individuals growing on the gravel and scree are shown. Ontogenesis of individuals of D. fragile is full and complex, consistig of ontogenesis of a seed individual and the reduced ontogeneses of affiliated individuals. Ontogenesis of seed individuals takes place in the sequence of the following phases of a morphogenesis: primary shoot (p,j) - primary bush (im) - bed (v,g1,g2,g3) - tillering particle (ss, s). The morphogenesis of individuals of a vegetative origin is characterized by the following phases: partial shoot (im) - partial shoots and bushes (v,g1,g2,g3, ss) - partial shoot (s). The course of an ontomorphogenesis of the seed and vegetative individuals growing on the scree is similar to ontogenesis of the individuals growing on the gravel. Habitat-dependent morphological and dynamic polyvariety of development of D. fragile individuals is pointed out.